Thanks Steve.
OK, this is what I did. Tested the tweeters - no short.
I decided to install both tweeters, and hey presto - they're working perfectly. I think you were right, that I must have shorted something the first time. Anyway, I'm very happy now everything's working. I should have changed the tweeters years ago - they are 25 years old, and now the speakers sound better than I can ever remember.
So for anyone thinking about replacing their tweeters - just do it, its really worth it. Its like having a new set of speakers again.
I used SEAS H519 25TFFC NEW 2015 Manufacture and no mods were necessary - they dropped straight in, albeit, because the flanges are not as deep, they are recessed a millimetre or two.
Best regards,
John